shapeNarrowly elongated behind eyes, 1.55 times as broad as long, in frontal view 1.25 as broad as high. Ocellar triangle 1.3 times shorter than OOL. POL 4 times as long as Od. Temples look elongated, about twice longer than LO. In frontal view eye twice as high as long, about as long as malar space. Face 1.5 as broad as high, 1.5 times as broad as height of eye, 4 times higher than clypeus. Maxillary palpi as long as height of face and clypeus combined [TL]. Head roundly narrowed behind eyes, 1.6 times as wide as long. Ocellar triangle width 1.3 times [shorter than ?] OOL; POL is 4 times the diameter of the ocellus. Temples twice as long as the transverse diameter of the eye. The longitudinal diameter of the eye is almost equal to the malar space. Face width 1.5 times its height. Maxillary palps equal to height of face with clypeus [T14].
sculptureHead densely puctate. Face very finely conversely striate. Vertex finely transversely striate. Temple finely longitudinally striate. Clypeus finely punctate, shining [TL]. The face with weak arcuate folds, the vertex behind the ocelli with transverse folds [T14].
colorPalpi dark brown [TL].
antenna
shape ♀
shape ♂Antennae with 20 segments, as long as body. Scape slightly widened, 1.5 times as long as wide; 3rd segment 3 times as long as wide, the others elongated and apical segments twice as long as wide [TL][T14].
colorThe basal part of antennae dark-brownish [TL].
mesosoma
shape1.5 times as long as high. Propodeum with a clear transverse carina and obtuse lateral tubercles. Propodeum anteriorly with a longitudinal carina ending at transverse carina with two small teeth. [TL].
sculptureMesonotum rugose-punctate and scutellum as well. Sides of mesonotum rugose without clear longitudinal elements. Propodeum rugulose punctate with granulate sculpture [TL].
colorProthorax partially dark-brownish [TL].
legs
shapeHind femur 3.5 times as long as broad. Outer and inner spurs of hind tibiae equal in length, as long as 1/2 of 1st metatarsal segment, 5th segment as long as 3rd, a little shorter than 2nd, 1.3 times longer than 4th [TL].
sculpture
colorLegs dark, coxae almost black; fore femora brown with lightly apex, tibiae brown-yellowish with pale basal part, tarsi light-brown; hind femora dark-brown, tibiae brown with pale basal part, tarsi light-brown [TL]. Legs mostly dark-colored, hind tibia brownish-yellow [T14].
wings
shape1-R1 0.7 as long as pterostigma. The marginal cell of the fore wing is twice as large as the 2nd submarginal cell. 3-SR vein 1.5 times shorter than r, 4 times shorter than SR1. 1-CU1 shorter than cu-a [TL].
colorWings faintly infuscate. Pterostigma and veins brown-yellowish [TL]. The wings are moderately darkened [T14].
metasoma
shapeCarapace gradually enlarged posteriorly, 1.7 times as long as maximal broad on its 1/3. In lateral view almost parallel-sided, about 1.2 times as high posteriorly as anteriorly [TL]. Metasoma widened towards apex, 1.4 times as long as wide [T14].
sculptureCarapace longitudinally rugulose, fine winding undulations ending at its 2/3, apex without longitudinal elements of sculpture [TL]. Metasoma with coarse and sinuous longitudinal folds reaching its apical third [T14].
colorCarapace and aperture partially dark-brownish [TL].
♀ ovipositor
♂ apertureAMA big, bean-formed; 0.7 as broad as carapace, about 0.5 height of carapace [TL]. The aperture is 0.7 times the width of the metasoma and half its height, bean-shaped [T14].
other info
distributionCZ [FE].
remarksDiagnosis: This species by some characters (antennae, head elongate behind eyes, shape of AMA, partially sculptured carapace) is close to M. temporalis. The comparatively short 3-SR, enlarged and almost parallel-sided carapace (laterally) of M. macrellips sp.n. are sufficient for delimitating these two species. Remarks: In the absence within series of M. temporalis of a similar tendency (shortened 3-SR, enlarged to apex carapace) arid, at the same time, the absence of series within M. macrellips sp.n. to discern the variation and, of course, the direction of variation, then it represents an independent species. Ultimately, such feature as carapace's shape (that is obviously different at a simple examination in both species) put these two species apart. Although, as we have seen in case of M. risorius with variable shapes of carapace within big series, then quite possible that it is only a variety of M. temporalis. Even short 3-SR (that is distinctly apparent on specimen M. macrellips sp.n. in comparison to comparatively long 3-SR within M. temporalis) can be just a part of variation (a similar case is described within M.foersteri's series) [TL].
hosts
figures
fig. 1: head in dorsal view (Tobias & Lozan 2003)
fig. 2: head in frontal view (Tobias & Lozan 2003)
M a l e T y p e m a t e r i a l e x a m i n e d : Holotype: ČR, Nový Jičin, 29.V.1930, Igt. Gregor ["macrellips"20], Id. E t y m o l o g y : This species is given a name macrellips that often is used on the labels of the Gregor's material.
D i m e n s i o n s : Body length 3.1mm.
H e a d : Narrowly elongated behind eyes, 1.55 times as broad as long, in frontal view 1.25 as broad as high. Ocellar triangle 1.3 times shorter than OOL. POL 4 times longer than Od. Temples look elongated, about twice longer than LO. In frontal view eye twice as high as long, about as long as malar space. Face 1.5 as broad as high, 1.5 times as broad as LD, 4 times higher than clypeus. Maxillary palpi as long as height of face and clypeus combined. Antennae with 20 segments, as long as body length. Scape slightly widened, 1.5 times as long as wide; Is' flagellum segment 3 times as long as wide, the others elongated and apical segments twice as long as wide.
M e s o s o m a : 1.5 times as long as high. Propodeum with a clear transverse carina and obtuse lateral tubercles.
W i n g s : Length of cell R along metacarp 0.7 as long as length of pterostigma. Cell R twice greater than Rm2. 3-SR vein 1.5 times shorter than r, 4 times shorter SRI. 1-CU1 shorter than cu-a.
L e g s : Hind femur 3.5 times as long as broad. Outer and inners spurs of hind tibiae equal in length, as long as 1/2 of 1SI metatarsal segment, 5Ih segment as long as 3rd, a little shorter than 2nd, 1.3 times longer than 4th.
M e t a s o m a : Carapace gradually enlarged posteriorly, 1.7 as long as maximal broad on its 1/3. In lateral view almost parallel-sided, about 1.2 times as high posteriorly as anteriorly. AMA big, bean-formed; 0.7 as broad as carapace, about 0.5 height of carapace.
S c u l p t u r e : Head dense puctate. Face very fine conversely striate. Vertex finelly transversely striate. Temple finely longitudinally striate. Clypeus finely punctate, shining. Mesonotum rugose-punctate and scutellum as well. Sides of mesonotum rugose without clear longitudinal elements. Propodeum rugulose punctate with granulate sculpture, anteriorly with a longitudinal carina ending to transverse carina and looking as two small teeth. Carapace longitudinally rugulose, fine winding undulations ending to its 2/3, apex without longitudinal elements of sculpture.
C o l o u r : Body black. Palpi dark brown. The basal part of antennae, prothorax, carapace and AMA partially dark-brownish. Legs dark, coxae almost black; fore femora brown with lightly apex, tibiae brown-yellowish with lightly basal part, tarsi light-brown; hind femora dark-brown, tibiae brown with lightly basal part, tarsi light-brown. Wings faintly infuscate. Pterostigma and veins brown-yellowish.
D i a g n o s i s : This species by some characters (antennae, head elongate behind eyes, shape of AMA, partially sculptured carapace) is closed to M. temporalis. The comparatively short 3-SR, enlarged carapace and almost parallel-sided carapace (laterally) within M. macrellips sp.n. are sufficient for delimitating these two species.
In the absence within series of M. temporalis of a similar tendency (shortened 3-SR, enlarged to apex carapace) arid, at the same time, the absence of series within M. macrellips sp.n. to discern the variation and, of course, the direction of variation, then it represents an independent species. Ultimately, such feature as carapace's shape (that is obviously different at a simple examination in both species) put these two species apart. Although, as we have seen in case of M. risorius with variable shapes of carapace within big series, then quite possible that it is only a variety of M. temporalis. Even short 3-SR (that is distinctly apparent on specimen M. macrellips sp.n. in comparison to comparatively long 3-SR within M. temporalis) can be just a part of variation (a similar case is described within M.foersteri's series).
D i s t r i b u t i o n : Czech Republic.
Data used in the multi-access key
These data are collected from the species descriptions and redescriptions, from published figures and photographs and from own measureements and observations. For a description of the different variables and value classes see this pdf.
morphology
basic information
female
male
body length (mm):
(2.7) 2.7 - 3.3 (3.5)
(2.7) 2.7 - 3.3 (3.5)
number of antennal segments:
16
19 - 21
head, dorsal view
Head width : length
very narrrow
very narrrow
Temples narrowing - widening
parallel
parallel
Eye length : temple length
<= 0.75
<= 0.75
Roundness temple
medium
medium
POL : OD
3 - 4, > 4
3 - 4, > 4
Ocellar triangle shape
very narrow
very narrow
Excavation of head
25 - 35%, > 35%
25 - 35%, > 35%
head, lateral view
Head height : length
---
---
Eye height : length
---
---
Temple length : eye length
---
---
head, frontal view
Head width : height
narrow, wide
narrow, wide
Face width : height
very narrow, narrow
very narrow, narrow
Eyes converging ventrally
distinctly diverging
distinctly diverging
EH : MSL
very short
very short
Clypeus width : height
semicircular, transverse
semicircular, transverse
Clypeus extending above bottom of eyes
far below
far below
Tentorial pits
very small
very small
Shape of lower margin clypeus
convex, truncate
convex, truncate
antennae
Antennal shape
---
filiform, not widened
Segment 3 length : width
---
medium
Penultimate segment length : width
---
long
Length of antenna : length of body
---
4: very long
mesosoma, lateral view
Length : height
medium
medium
transverse carina
well developed
well developed
lateral teeth
present
present
mesosoma, ventral view
Prepectal carina lamelliform
indistinct
indistinct
legs
Hind femur length : width ratio
medium
medium
Hind tibia length : hind tarsus length ratio
---
---
Hind tibia (longest) spur length : metatars length ratio
long
long
wings
Vein 1-R1 length : pterostigma length
medium
medium
Vein r length : vein 3-SR length
short, long
short, long
Shape vein SR1 (straight / concave / convex)
straight or S-shaped
straight or S-shaped
S-shape vein SR1
not S-shaped
not S-shaped
cell M : cell Sm2
Sm2 medium, Sm2 small
Sm2 medium, Sm2 small
Pterostigma length : width ratio
short
short
Angle between radial vein (r) : vein 1-R1
medium, blunt
medium, blunt
metasoma, dorsal view
Length : width ratio of metasoma
short, medium
short, medium
Shape of metasoma
tapering to base
tapering to base
metasoma, lateral view
Length : height ratio of metasoma
medium
medium
posterior : anterior height ratio of metasoma
small, medium
small, medium
Position of maximum height
medium-posterior
medium-posterior, posterior
metasoma, ventral view
Metasoma posteriorly curved forwards
---
---
male apical aperture
Aperture width : metasoma width
no aperture
medium, large
Aperture shape
n. a.
(nearly) round, oval
Median process shape
n. a.
horizontal oval
sculpture
head
general coarseness
fine
fine
clypeus
coarseness punctation
punctulate
punctulate
density punctation
---
---
face
sculpture type
striate
striate
sculpture direction
transverse, diagonal/concentric
transverse, diagonal/concentric
vertex and temple
sculpture type
striate (striolate)
striate (striolate)
mesonotum
sculpture type
---
---
scutellum
middle
punctate, rugose / striate
punctate, rugose / striate
lateral
punctate, rugose / striate
punctate, rugose / striate
metasoma
base - mid sculpture
longitudinally rugose / rugose-striate
longitudinally rugose / rugose-striate
sculpture at apex
rugulose
rugulose
coarse / fine boundary
mid (50 - 70%)
mid (50 - 70%)
color
head
mandibles color pattern
---
---
palpi, lightest color
black, dark brown
black, dark brown
antennae
scapus, lightest color
black, dark brown
black, dark brown
segment 3 and 4, lightest color
black, dark brown
black, dark brown
legs
fore coxae lightest colour
dark
dark
fore femur lightest colour
medium light
medium light
fore femur color pattern
tip light
tip light
hind femur lightest colour
dark
dark
hind femur color pattern
entirely dark
entirely dark
hind tibia lightest colour
medium light
medium light
hind tibia color pattern
narrow light base, wide light base
narrow light base, wide light base
hind basitarsus lightest colour
medium light
medium light
hind tarsus segment 2-3 lightest colour
medium light
medium light
wings
middle veins lightest colour
medium light
medium light
basal veins lightest colour
medium light, light
medium light, light
metasoma
metasoma colour pattern
entirely dark
entirely dark
metasoma lightest colour
dark
dark
metasoma base, colour pattern
entirely dark
entirely dark
metasoma base, light spots
n.a.
n.a.
metasoma base, light band
n.a., regular posterior edge
n.a., regular posterior edge
Chelonus macrellips in published keys
TL: Tobias & Lozan 2003
Central-European species of Microchelonus Szépligeti (Hymenoptera, Braconidae) with very big apical metasomal aperture of males.
T14: Tobias, 2010
Palearctic species of Microchelonus Szépligeti (Hymenoptera: Braconidae, Cheloninae): key to species.